They are sentient, yet there are times when they are piloted by Shadow Angels. The Shadow Angels facilitate their harvesting via giant, floating harvesting machines called harvest beasts which are guarded by giant machines called Cherubim Soldiers. The Shadow Angels began invading human cites on Earth to Harvest human beings like cattle, dubbing them "wingless ones" (羽なし hanenashi), to extract the Prana from the harvested humans not only to serve as energy and nutrition to them, but also to feed the legendary Tree of Life. Along with the disaster came the return of the Shadow Angels, who have been in slumber in their city of Atlandia since their battle with Apollonius. Apollonius joined forces with the humans to free them from their oppression, by using the legendary giant fighting robot Aquarion.Įleven years prior to the story in the Holy Genesis Era, a disaster only referred to as the " Great Catastrophe" occurred, where Earth's magnetic field shifted, destroying significant parts of the world and killing off many of its inhabitants. That is, until one of their kind, Apollonius, fell in love with a female human warrior, Celiane.
Twelve thousand years in the past, mankind was at mercy of the mythical creatures known as Shadow Angels, immortal winged beings with overwhelming powers and technology.
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